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INVERTEBRATES
It is virtually impossible
to be an expert in the taxonomy of all of the wildlife in an area like
the one in this study. This becomes clearly evident as one attempts to identify
the great variety of invertebrates in the area. Although several
good general references are available and have been used in this study,
(Pennak, 1978, Needham and Needham, 1962), it is difficult
to find references that are broad enough to cover the variety of organisms
found yet specific enough to the area. Hopefully, reliable data concerning
the invertebrates of the area will accumulate as the study continues. The
following is a brief review of several categories for which some useful
references have been found. Is is important to recognize that it is not the
result of any systematic studies of the invertebrates in the area.
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Crustaceans: The
Wisconsin Department of Natural
Resources has done many studies of the freshwater crustaceans of Wisconsin.
One such study (Torke, 1979) presents crustacean
zooplankton data for 190 selected Wisconsin inland lakes sampled in 1973
and 1974. Although the study includes no Crawford County lakes, the report
may serve to suggest species that are likely to occur in the area. Taxonomic
keys to the copepods of Wisconsin are available to those who would accept
the challenge of collecting and identifying the organisms of the oxbow lakes
in the study area.
Joan Jass, of the Milwaukee Public Museum, has studied
the fairy shrimp of Wisconsin. At the 6th. annual wetland scientists forum,
sponsored by the Wisconsin Wetlands Association,
she reported on studies that elaborate three species for the state. No
fairy shrimp have yet been found during my studies in this area.
A key to the 11 adult terrestrial isopods thus far
documented for Wisconsin can be found in Jass and
Klausmeier (1996). I have no data regarding these organisms for Crawford
County at this time.
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Diplopods:
Coming from the southeast part of the state, one cannot but be impressed
by the differences in the millipedes of this area. Although I have not
located taxonomic keys for these animals, a review of the various species
can be found in Watermolen, (1995). In
Wisconsin Millipeds: A Preliminary Species List and
a Catalog of Records, the author lists 26 species representing 13 families
in 6 orders for the state. These numbers are based on reviews of milliped
literature and examination of specimens from various museum collections.
Crawford County is represented by seven (7) species from six different
families. Another two (2) species representing two more families are listed
for the general area without specific reference to the county. Ten other
species listed as "potential Wisconsin species" because they have been found
in neighboring states and have habitat needs likely to occur in Wisconsin
were also described in the paper. At this time, given my lack of reliable
taxonomic literature, I believe that I have recognized at least two (2)
species in the study area. One is Narceus americanus
anularis. Recently, I found a specimen that measured a full 10cm. in
length and that I believed to be this species. The other is Pleuroloma
flavipes. Both of these species, are documented for Crawford County
in the Watermolen reference.
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Arachnids: Arachnids
are plentiful in the study area. To date, no systematic efforts have been
made on any of them except the hard ticks (family - Ixodidae). These are
both plentiful and bothersome, especially in the spring of the year. Keirans and Litwak (1989) present a pictorial
key to the adults of hard ticks east of the Mississippi River. I have used
the key to identify adult ticks to the genus Ixodes in the area but have
not attempted identification to species. Such identification would
require a very good binocular microscope. A variety of ticks from the area
have been preserved for future taxonomic efforts.
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Insects: The following
is a description of the references that have been found for several categories
of both aquatic and terrestrial insects that have attracted attention in
the study area.
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Mosquitos (Order - Diptera; Family - Culicidae)
are very abundant in the study area, especially early in the spring and
during periods following heavy rains. Huge densities of mosquito larvae
can be seen in the floodplain soon after the spring thaw. Early attempts
to survey the mosquitos of Wisconsin (Dickinson,
1944) do not include records from Crawford County. Taxonomic keys offered
by this reference have undoubtedly been improved upon by later publications.
Following scares regarding the LaCrosse strain of encephalitis in the mid
to late 1970's, the professional health literature became a good source
of many illustrated keys to assist health professionals in their efforts
to identify Aedes triseriatus, the common vector
of the disease. As a temporary health professional with Waukesha County at
that time, I became exposed to a variety of such keys -- especially those
of Chester J. Stojanovich and Harry Pratt. Studies done by the Division of
Allied Health of the University of Wisconsin - Eau Claire should be helpful
to those interested in Wisconsin mosquito taxonomy.
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A variety of publications of the Wisconsin Department
of Natural Resources are helpful for identifying aquatic invertebrates, especially
insects, of Wisconsin waters. Aquatic Insects of Wisconsin (Hillsenhoff, 1995) provides taxonomic keys
to the genera of aquatic insects of the orders ephemeroptera (mayflies),
odonata (dragonflies), plecoptera (stoneflies), tricoptera (caddisflies),
megaloptera (alderflies and fishflies), heteroptera (aquatic bugs), lepidoptera
(aquatic moths) neuroptera (spongilliflies), coleoptera (aquatic beetles)
and diptera (aquatic flies and midges). Keys for both larval and adults
are included when these are aquatic forms. The keys are thoroughly illustrated
and an extensive glossary is included in the publication although no geographical
data are included.
In an earlier publication (Hillsenhoff, 1995b), Using a Biotic Index
to Evaluate Water Quality in Streams, the Wisconsin DNR includes keys
to the species of 22 genera of aquatic insects that are useful in assessing
water quality.
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The Northern
Prairie Wildlife Research Center offers a number of online references
that are specific to counties of the various states. The center lists only
10 moth species from 2 families and 16 species of butterflies representing
5 families for Crawford County. Nice photographs of both adult and immature
stages of the butterflies and moths are available online. Luckily, more
specific literature is available for the butterflies. Swengel (1991), is a handy reference showing
the species of butterflies found in southwestern Wisconsin along with their
status, occurrence and habitat. It has graphic representations of the seasonal
occurrence for each species. The reference lists 111 species for southwestern
Wisconsin but does not break the data down by county. To date, without much
effort devoted to the group, I have identified 18 butterflies in the study
area. The tentative identifications are shown below. The familiar "woolly
bear" caterpillar of the Isabella moth (probably Isea isabella) and
one of the prominents (possibly Datana ministra), recognized by its
habit of holding its tail erect, have been recognized among the moths. At
this time no specimens have been collected for more reliable taxonomy. This
should be a point of further study.
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Butterflies of the Study Area
Giant Swallowtail - Heraclides cresphontes
Eastern Black Swallowtail - Papilio polyxenes
Eastern Tiger Swallowtail - Pterourus glaucus
Cabbage White - Artogeia rapae
Common (Clouded) Sulfur - Colias philodice
Great Spangled Fritillary - Speyeria cybele
Meadow Fritillary - Clossiana bellona
Pearly Crescentspot - Phyciodes tharos
Hop Merchant - Polygonia comma
Mourning Cloak - Nymphalis antiopa
Milbert's Tortoiseshell - Aglais milberti
American Painted Lady - Vanessa cardui
Red Admiral - Vanessa atalanta
Red-spotted Purple - Basilarchia arthemis
Viceroy - Basilarchia archippus
Hackberry Butterfly - Asterocampa celtis
Monarch - Danaus plexippus
Silver-spotted Skipper - Epargyrus clarus
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The dragonflies is another insect group for
which there is a good specific reference. Common Dragonflies of Wisconsin
(Legler et. al., 1998) is a nicely illustrated
reference for identifying these insects as well as a few of Wisconsin's
damselflies. The book includes range maps and seasonal distribution graphs
as well as useful descriptive material for each species. The authors list
110 species for the state. Examination of the range maps suggests that 34
species have been documented in Crawford County. This includes 6 darners
(Family Aeshnidae), 6 clubtails (Family Gomphidae), 1 spiketail (Family
Cordulegastridae), 1 cruiser (family Macromiidae) 3 emeralds (Family Corduliidae)
and 15 skimmers (Family Libellulidae) among the dragonflies and 2 damselflies.
Many of the photographs which illustrate this book are those of David Westover,
a nature photographer who has worked with the Wisconsin chapter of the Nature Conservancy. His photographs of
Wisconsin dragonflies can be found online at Digital Dragonflies. This website also has
many photos of butterflies and damselflies from the Texas area. In addition
to this reference, online images of many of the local species can be found
at the websites for the Dragonflies and
Damselflies of Massachusetts.or the Michigan
Odonata Survey. To date, the dot-tailed whitetail (Leucorrhinia
intacta), common whitetail (Plathemis lydia), common green darner
(Anax junius), band-winged meadowhawk (Sympetrium semicinctum),
and ruby meadowhawk (Sympetrium rubicundulum) have been tentatively
identified in the study area. Among the damselflies I've recognized the
ebony jewelwing (Calopteryx maculata) as well as members of the
spreadwing damselflies and bluets.
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